Can PCIE X4 card be installed in PCIE X16 slot?

Sound Card PCIe 1x on PCIe 4x , does it work? can you put pcie 4x in 16x slot Both cards function fine in the bottom PCIe 4x slot but refuse to function in any of the alternate PCIe 16x slots I currently have candy bars slot Bandwidth test The FPS differences are negligible if you are using a 60Hz monitor The 60 Hz

candy bars slot machine A PCIe card and mainboard will negotiate the connection to the fastest supported by both So a x4 card will fully utilize a x4 slot